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11.1HOW SKELETAL MUSCLES PRODUCE MOVEMENTS OBJECTIVES •Describe the relationship between bones and skeletal muscles in producing body movements. •Define lever and fulcrum, and compare the three types of levers based on location of the fulcrum, effort, and load. •Identify the types of fascicle arrangements in a skeletal muscle, and relate the arrangements to strength of contraction and range of motion. •Explain how the prime mover, antagonist, synergist, and fixator in a muscle group work together to produce movement. Muscle Attachment Sites: Origin and Insertion Those skeletal muscles that produce movements do so by exerting force on tendons, which in turn pull on bones or other structures (such as skin). Most muscles cross at least one joint and are usually attached to articulating bones that form the joint (Figure11.1a). Figure11.1 Relationship of skeletal muscles to bones. Muscles are attached to bones by tendons at their origin and insertion. Skeletal muscles produce movements by pulling on bones. Bones serve as levers, and joints act as fulcrums for the levers. Here the lever–fulcrum principle is illustrated by the movement of the forearm. Note where the load (resistance) and effort are applied in (b). In the limbs, the origin of a muscle is usually proximal and the insertion is usually distal. John Gibb/Imagineering. Where is the belly of the muscle that extends the forearm located? When a skeletal muscle contracts, it pulls one of the articulating bones toward the other. The two articulating bones usually do not move equally in response to contraction. One bone remains stationary or near its original position, either because other muscles stabilize that bone by contracting and pulling it in the opposite direction
or because its structure makes it less movable. Ordinarily, the attachment of a muscle's tendon to the stationary bone is called theorigin(OR i jin); the attachment of the muscle's other tendon to the movable bone is called‐ ‐ theinsertion(in SER shun). A good analogy is a spring on a door. In this example, the part of the spring‐‐ attached to the frame is the origin; the part attached to the door represents the insertion. A useful rule of thumb is that the origin is usually proximal and the insertion distal; the insertion is usually pulled toward the origin. The fleshy portion of the muscle between the tendons is called thebelly (body), the coiled middle portion of the spring in our example. Theactionsof a muscle are the main movements that occur when the muscle contracts. In our spring example, this would be the closing of the door. Certain muscles are also capable ofreverse muscle action (RMA). This means that during specific movements of the body the actions are reversed and therefore the positions of the origin and insertion of a specific muscle are switched. Muscles that move a body part often do not cover the moving part. Figure11.1b shows that although one of the functions of the biceps brachii muscle is to move the forearm, the belly of the muscle lies over the humerus, not over the forearm. You will also see that muscles that cross two joints, such as the rectus femoris and sartorius of the thigh, have more complex actions than muscles that cross only one joint. Examples Anatomy Overview: Skeletal Muscle Anatomy Overview: The Muscular System‐Cross Section of Skeletal Muscle Lever Systems and Leverage In producing movement, bones act as levers, and joints function as the fulcrums of these levers. Aleveris a rigid structure that can move around a fixed point called afulcrum, symbolized by. A lever is acted on at two different points by two different forces: theeffort(E), which causes movement, and theload orresistance, which opposes movement. The effort is the force exerted by muscular contraction; the load is typically the weight of the body part that is moved or some resistance that the moving body part is trying to overcome (such as the weight of a book you might be picking up). Motion occurs when the effort applied to the bone at the insertion exceeds the load. Consider the biceps brachii flexing the forearm at the elbow as an object is lifted (Figure11.1b). When the forearm is raised, the elbow is the fulcrum. The weight of the forearm plus the weight of the object in the hand is the load. The force of contraction of the biceps brachii pulling the forearm up is the effort. The relative distance between the fulcrum and load and the point at which the effort is applied determine whether a given lever operates at a mechanical advantage or a mechanical disadvantage. For example, if the load is closer to the fulcrum and the effort farther from the fulcrum, then only a relatively small effort is required to move a large load over a small distance. This is called amechanical advantage. If, instead, the load is farther from the fulcrum and the effort is applied closer to the fulcrum, then a relatively large effort is required to move a small load (but at greater speed). This is called amechanical disadvantage. Compare chewing something hard (the load) with your front teeth and the teeth in the back of your mouth. It is much easier to crush the hard food item with the back teeth because they are closer to the fulcrum (the jaw or temporomandibular joint) than are the front teeth. Here is one more example you can try. Straighten out a paper clip. Now get a pair of scissors and try to cut the paper clip with the tip of the scissors (mechanical disadvantage) versus near the pivot point of the scissors (mechanical advantage). Levers are categorized into three types according to the positions of the fulcrum, the effort, and the load: 1.The fulcrum is between the effort and the load infirst class levers‐(Figure11.2a). (Think EFL.) Scissors and seesaws are examples of first class levers. A first class lever can produce either a‐‐
mechanical advantage or mechanical disadvantage depending on whether the effort or the load is closer to the fulcrum. (Think of an adult and a child on a seesaw.) As we've seen in the preceding examples, if the effort (child) is farther from the fulcrum than the load (adult), a heavy load can be moved, but not very far or fast. If the effort is closer to the fulcrum than the load, only a lighter load can be moved, but it moves far and fast. There are few first class levers in the body. One example is the lever formed by‐ the head resting on the vertebral column (Figure11.2a). When the head is raised, the contraction of the posterior neck muscles provides the effort E, the joint between the atlas and the occipital bone (atlanto‐ occipital joint) forms the fulcrum, and the weight of the anterior portion of the skull is the load. 2.The load is between the fulcrum and the effort insecond class levers‐(Figure11.2b). (Think ELF.) Second class levers operate like a wheelbarrow. They always produce a mechanical advantage because‐ the load is always closer to the fulcrum than the effort. This arrangement sacrifices speed and range of motion for force; this type of lever produces the most force. This class of lever is uncommon in the human body. An example is standing up on your toes. The fulcrumis the ball of the foot. The load is the weight of the body. The effort (E) is the contraction of the muscles of the calf, which raise the heel off the ground. 3.The effort is between the fulcrum and the load inthird class levers‐(Figure11.2c). (Think FEL.) These levers operate like a pair of forceps and are the most common levers in the body. Third class levers‐ always produce a mechanical disadvantage because the effort is always closer to the fulcrum than the load. In the body, this arrangement favors speed and range of motion over force. The elbow joint, the biceps brachii muscle, and the bones of the arm and forearm are one example of a third class lever‐ (Figure11.2c). As we have seen, in flexing the forearm at the elbow, the elbow joint is the fulcrum, the contraction of the biceps brachii muscle provides the effort E, and the weight of the hand and forearm is the load. Examples Animation: Contraction and Movement Effects of Fascicle Arrangement Recall from Chapter10that the skeletal muscle fibers (cells) within a muscle are arranged in bundles known asfascicles(FAS i kuls). Within a fascicle, all muscle fibers are parallel to one another. The fascicles,‐ ‐ however, may form one of five patterns with respect to the tendons: parallel, fusiform (spindle shaped, narrow‐ toward the ends and wide in the middle), circular, triangular, or pennate (shaped like a feather) (Table11.1). Table11.1Arrangement of Fascicles PARALLEL Fascicles parallel to longitudinal axis of muscle; terminate at either end in flat tendons.
Example:Sternohyoid muscle (see Figure11.8a) FUSIFORM Fascicles nearly parallel to longitudinal axis of muscle; terminate in flat tendons; muscle tapers toward tendons, where diameter is less than at belly. Example:Digastric muscle (see Figure11.8a) CIRCULAR Fascicles in concentric circular arrangements form sphincter muscles that enclose an orifice (opening). Example:Orbicularis oculi muscle (see Figure11.4a) TRIANGULAR Fascicles spread over broad area converge at thick central tendon; gives muscle a triangular appearance. Example:Pectoralis major muscle (see Figure11.3a) PENNATE Short fascicles in relation to total muscle length; tendon extends nearly entire length of muscle. UNIPENNATE Fascicles arranged on only one side of tendon. Example:Extensor digitorum longus muscle (see Figure11.22b) BIPENNATE Fascicles arranged on both sides of centrally positioned tendons. Example:Rectus femoris muscle (see Figure11.20a) MULTIPENNATE Fascicles attach obliquely from many directions to several tendons.
Example:Deltoid muscle (see Figure11.10a) Fascicular arrangement affects a muscle's power and range of motion. As a muscle fiber contracts, it shortens to about 70% of its resting length. The longer the fibers in a muscle, the greater the range of motion it can produce. However, the power of a muscle depends not on length but on its total cross sectional area, because a‐ short fiber can contract as forcefully as a long one. So the more fibers per unit of cross sectional area a muscle‐ has, the more power it can produce. Fascicular arrangement often represents a compromise between power and range of motion. Pennate muscles, for instance, have a large number of short fibered fascicles distributed over‐ their tendons, giving them greater power but a smaller range of motion. In contrast, parallel muscles have comparatively fewer fascicles, but they have long fibers that extend the length of the muscle, so they have a greater range of motion but less power. CLINICAL CONNECTIONIntramuscular Injections Anintramuscular (IM) injectionpenetrates the skin and subcutaneous layer to enter the muscle itself. intramuscular injections are preferred when prompt absorption is desired, when larger doses than can be given subcutaneously are indicated, or when the drug is too irritating to give subcutaneously. the common sites for intramuscular injections include the gluteus medius muscle of the buttock (see Figure11.3b), lateral side of the thigh in the midportion of the vastus lateralis muscle (see Figure11.3a), and the deltoid muscle of the shoulder (see Figure11.3b). muscles in these areas, especially the gluteal muscles in the buttock, are fairly thick, and absorption is promoted by their extensive blood supply. to avoid injury, intramuscular injections are given deep within the muscle, away from major nerves and blood vessels. intramuscular injections have a faster speed of delivery than oral medications but are slower than intravenous infusions. Coordination among Muscles Movements often are the result of several skeletal muscles acting as a group. Most skeletal muscles are arranged in opposing (antagonistic) pairs at joints—that is, flexors–extensors, abductors–adductors, and so on. Within opposing pairs, one muscle, called theprime moveroragonist(= leader), contracts to cause an action while the other muscle, theantagonist(anti= against), stretches and yields to the effects of the prime mover.‐ In the process of flexing the forearm at the elbow, for instance, the biceps brachii is the prime mover, and the triceps brachii is the antagonist (see Figure11.1a). The antagonist and prime mover are usually located on opposite sides of the bone or joint, as is the case in this example. With an opposing pair of muscles, the roles of the prime mover and antagonist can switch for different movements. For example, while extending the forearm at the elbow against resistance (i.e., lowering the load shown in Figure11.2c), the triceps brachii becomes the prime mover, and the biceps brachii is the antagonist. If a prime mover and its antagonist contract at the same time with equal force, there will be no movement. Figure11.2Lever structure and types of levers. Levers are divided into three types based on the placement of the fulcrum, effort, and load (resistance).John Gibb/Imagineering. Which type of lever produces the most force?
Sometimes a prime mover crosses other joints before it reaches the joint at which its primary action occurs. The biceps brachii, for example, spans both the shoulder and elbow joints, with primary action on the forearm. To prevent unwanted movements at intermediate joints or to otherwise aid the movement of the prime mover, muscles calledsynergists(SIN er jists;‐‐syn= together;‐‐ergon= work) contract and stabilize the intermediate joints. As an example, muscles that flex the fingers (prime movers) cross the intercarpal and radiocarpal joints (intermediate joints). If movement at these intermediate joints were unrestrained, you would not be able to flex your fingers without flexing the wrist at the same time. Synergistic contraction of the wrist extensor muscles stabilizes the wrist joint and prevents unwanted movement, while the flexor muscles of the fingers contract to bring about the primary action, efficient flexion of the fingers. Synergists are usually located close to the prime mover. Some muscles in a group also act asfixators, stabilizing the origin of the prime mover so that the prime mover can act more efficiently. Fixators steady the proximal end of a limb while movements occur at the distal end. For example, the scapula is a freely movable bone that serves as the origin for several muscles that move the arm. When the arm muscles contract, the scapula must be held steady. In abduction of the arm, the deltoid muscle serves as the prime mover, and fixators (pectoralis minor, trapezius, subclavius, serratus anterior muscles, and others) hold the scapula firmly against the back of the chest (see Figure11.14a, b). The insertion of the deltoid muscle pulls on the humerus to abduct the arm. Under different conditions—that is, for different movements—and at different times, many muscles may act as prime movers, antagonists, synergists, or fixators. In the limbs, acompartmentis a group of skeletal muscles, their associated blood vessels, and associated nerves, all of which have a common function. In the upper limbs, for example, flexor compartment muscles are anterior, and extensor compartment muscles are posterior. CLINICAL CONNECTIONBenefits of Stretching The overall goal ofstretchingis to achieve normal range of motion of joints and mobility of soft tissues surrounding the joints. For most individuals, the best stretching routine involvesstatic stretching, that is, slow sustained stretching that holds a muscle in a lengthened position. The muscles should be stretched to the point of slight discomfort (not pain) and held for about 30 seconds. Stretching should be done after warming up to increase the range of motion most effectively. 1.Improved physical performance.A flexible joint has the ability to move through a greater range of motion, which improves performance. 2.Decreased risk of injury.Stretching decreases resistance in various soft tissues so there is less likelihood of exceeding maximum tissue extensibility during an activity (i.e., injuring the soft tissues). 3.Reduced muscle soreness.Stretching can reduce some of the muscle soreness that results after exercise. 4.Improved posture.Poor posture results from improper position of various parts of the body and the effects of gravity over a number of years. stretching can help realign soft tissues to improve and maintain good posture.
11.2HOW SKELETAL MUSCLES ARE NAMED OBJECTIVE •Explain seven features used in naming skeletal muscles. The names of most of the skeletal muscles contain combinations of the word roots of their distinctive features. This works two ways. You can learn the names of muscles by remembering the terms that refer to muscle features, such as the pattern of the muscle's fascicles; the size, shape, action, number of origins, and location of the muscle; and the sites of origin and insertion of the muscle. Knowing the names of a muscle will then give you clues about its features. Study Table11.2to become familiar with the terms used in muscle names. Table11.2Characteristics Used to Name Muscles NAMEMEANINGEXAMPLEFIGURE DIRECTION: ORIENTATION OF MUSCLE FASCICLES RELATIVE TO THE BODY'S MIDLINE RectusParallel to midlineRectus abdominis11.10c TransversePerpendicular to midlineTransversus abdominis11.10c ObliqueDiagonal to midlineExternal oblique11.10a SIZE: RELATIVE SIZE OF THE MUSCLE MaximusLargestGluteus maximus11.20c MinimusSmallestGluteus minimus11.20d LongusLongAdductor longus11.20a BrevisShortAdductor brevis11.20b LatissimusWidestLatissimus dorsi11.15b LongissimusLongestLongissimus capitis11.19a MagnusLargeAdductor magnus11.20b MajorLargerPectoralis major11.10a MinorSmallerPectoralis minor11.14a VastusHugeVastus lateralis11.20a SHAPE: RELATIVE SHAPE OF THE MUSCLE
DeltoidTriangularDeltoid11.15b TrapeziusTrapezoidTrapezius11.3b SerratusSaw toothed‐Serratus naterior11.14b RhomboidDiamond shaped‐Rhomboid amjor11.15d OrbicularisCircularOrbicularis couli11.4a PectinateComblikePectineus11.20a PiriformisPear shaped‐Piriformis11.20d PlatysFlatPlatysma11.4c QuadratusSquare, four sided‐Quadratus efmoris11.20d GracilisSlenderGracilis11.20a ACTION: PRINCIPAL ACTION OF THE MUSCLE FlexorDecreases joint angleFlexor carpi radialis11.17a ExtensorIncreases joint angleExtensor carpi ulnaris11.17d AbductorMoves bone away from midlineAbductor pollicis longus11.17e AdductorMoves bone closer to midlineAdductor longus11.20a LevatorRaises or elevates body partLevator scapulae11.14a DepressorLowers or depresses body partDepressor labii inferioris11.4b SupinatorTurns palm anteriorlySupinator11.17c PronatorTurns palm posteriorlyPronator teres11.17a SphincterDecreases size of an openingExternal anal sphincter11.12 TensorMakes body part rigidTensor fasciae latae11.20a RotatorRotates bone around longitudinal axisRotatore11.19b NUMBER OF ORIGINS: NUMBER OF TENDONS OF ORIGIN BicepsTwo originsBiceps brachii11.16a TricepsThree originsTriceps brachii11.16b QuadricepsFour originsQuadriceps femoris11.20a
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