While the striatum basal carries on to generate new cells, keratinocytes present in the striatum spinosum are moved or embedded in the stratum granulosum. The nucleus and other cellular organelles disintegrate when the cells die, leaving keratinocytes, or the cell membrane behind, forming the stratum corneum and stratum lucidum. J., Birarda, G., Auerbach, A., Koskinen, K., Wolf, P., & Holman, H.